Hargreaves lab
Publications by theme
some papers are listed in >1 category. chronological order here

Some Lab Favourites blue = student in Hargreaves lab
Urbanization weakens the global latitudinal diversity gradient in birds.
Griffith, Sunday, AL Hargreaves. 2026.
PNAS. published online Aug 18 10.1073/pnas.2527860123
Biotic interactions are more often important at species' warm vs. cool range-edges
Paquette, Hargreaves. 2021.
Ecology Letters. 24(11): 2427-2438
Hargreaves, Samis, Eckert. 2014.
American Naturalist. 183: 175-183.
Latitudinal gradients in predation persist in urban environments
Hargreaves AL, J Ensing, O Rahn, F Oliviera, J Burkiewicz & 28 co-authors. 2024.
Nature Ecology & Evolution. 8: 1897-1906.
(more latitudinal gradients stuff here)
Mapping nationally and globally at-risk species to identify hotspots for (and gaps in) conservation
Hardouin, Hargreaves. 2023.
Proceedings of the Royal Society B. 290 (1995), 20222307.
(more species at risk stuff here)



Species distributions & range limits
blue = student in Hargreaves lab
Shifting, expanding, or contracting? Range movement consequences for biodiversity
JF Brodie, BG Freeman, PD Mannion, AL Hargreaves. 2025.
Trends in Ecology & Evolution. 40: 439-448.
Temperate species underfill their tropical thermal potentials on land
Moore, NA, I Morales-Castilla, AL Hargreaves, 13 co authors, JR Bernhardt, JM Sunday. 2023.
Nature Ecology Evolution. 7: 1993-2003.
The evolution of plasticity across species ranges
Usui, T, D Lerner, IM Eckert, A Angert*, CJ Garroway*, AL Hargreaves*, L Lancaster*, JP Lessard*, F Riva*, C Schmidt*, K van der Burg*, KE Marshall. *order alphabetical. 2023.
Trends in Ecology & Evolution. 38(9) 831-842
Effects of species interactions on the potential for evolution at species' range limits
JM Alexander, DZ Atwater, RI Colautti, AL Hargreaves. 2022
Philosophical Transactions of the Royal Society B, 377 (1848): 20210020
Does pollen limitation limit plant ranges? Evidence and implications
Dawson-Glass, E, AL Hargreaves. 2022
Philosophical Transactions of the Royal Society B, 377: 20210014.
Haider, S, J Lembrechts, K McDougall, A Pauchard, 51 authors listed alphabetically including AL Hargreaves, T Seipel. 2022
Ecology & Evolution. 12(2): e8590.
Biotic interactions are more often important at species' warm vs. cool range-edges
Paquette, A, AL Hargreaves. 2021
Ecology Letters. 24(11): 2427-2438
Bontrager, M, T Usui, JA Lee-Yaw, DN Anstett, HA Branch, AL Hargreaves, CD Muir, AL Angert. 2021.
Evolution. 75: 1316-1333
High conservation priority of range-edge plant populations not matched by habitat protection or research effort. Caissy, P, S Klemet-N’Guessan, R Jackewi, CG Eckert, AL Hargreaves. 2020.
Biological Conservation. 249: 108732 (data & Rcode)
Spatial distribution and conservation hotspots of mammals in Canada
Cameron, V, AL Hargreaves. 2020.
FACETS. 5: 692-703
Miniaturizing species distributions to understand evolving range dynamics
Larsen, C, AL Hargreaves. 2020.
Ecography (short listed for E4 award).
Local adaptation primes cold-edge populations for range expansion but not warming-induced range shifts Hargreaves, AL & CG Eckert. 2019.
Ecology Letters. 22: 78-88. (data & R code, media coverage)
Expanding, shifting, and shrinking: The impact of global warming on species' elevational distributions Freeman B, J Lee-Yaw, J Sunday, AL Hargreaves. 2018.
Global Ecology & Biogeography. 27: 1268-1276. (media coverage)
Local maladaptation reduces expansion load during range expansion
Gilbert KJ, NP Sharp, AL Angert*, GL Conte*, JA Draghi*, F Guillaume*, AL Hargreaves*, R Matthey-Doret*, MC Whitlock. 2017. *order alphabetical
American Naturalist. 189: 368-380. (DOI, summary)
Hargreaves, AL, SF Bailey, RA Laird. 2015.
Journal of Evolutionary Biology. 28: 1489-1501. (Appendix, data & R code)
Hargreaves, AL, KE Samis, CG Eckert. 2014.
American Naturalist. 183: 175-183. (blog summary, DOI, Appendices, data)
Hargreaves, AL & CG Eckert. 2014.


Latitudinal & elevational patterns in biotic interactions
Urbanization weakens the global latitudinal diversity gradient in birds.
Griffith, J, J Sunday, AL Hargreaves. 2026.
PNAS. published online Aug 18 10.1073/pnas.2527860123
Geographic gradients in species interactions: from latitudinal patterns to ecological mechanisms.
Hargreaves, AL. 2024.
Annual Review of Ecology, Evolution, and Systematics. 55: 369-393.
Latitudinal gradients in predation persist in urban environments
Hargreaves AL, J Ensing, O Rahn, F Oliviera, J Burkiewicz, J Lafond, S Haeussler, B Best, K Lazda, H Slinn, E Martin, M Carlson, T Sformo, E Dawson-Glass, M Chuiffo, Y Vargas-Rodriguez, C García-Jiménez, I Gomes, S Klemet-N’Guessan, L Paolucci, S Joly, K Mehltreter, J Muñoz, C Buono, J Brodie, A Rodriguez-Campbell, T Veen, B Freeman, J Lee-Yaw, J Camilo Muñoz, A Paquette, J Butler, E Suaréz. 2024.
Nature Ecology & Evolution. 8: 1897-1906.
Clay larvae do not accurately measure biogeographic patterns in predation
Rodriguez-Campbell A, O Rahn, M Chuiffo, AL Hargreaves. 2024.
Journal of Biogeography. 51:1004-1013.
Gradients in how long seeds take to germinate could alter global patterns in predation strength
Martin, E, AL Hargreaves. 2023.
Journal of Biogeography. 50: 884-896.
Does pollen limitation limit plant ranges? Evidence and implications
Dawson-Glass, E, AL Hargreaves. 2021.
Philosophical Transactions of the Royal Society B, 377: 20210014.
Biotic interactions are more often important at species' warm vs. cool range-edges
Paquette, A, AL Hargreaves. 2021
Ecology Letters. 24(11): 2427-2438
Local adaptation to biotic interactions: a meta-analysis across latitudes
Hargreaves, AL, R Germain, M Bontrager, J Persi, A Angert. 2020.
American Naturalist. 195: 395-411 (press release)
Seed predation increases from the Arctic to the Equator and from high to low elevations
Hargreaves, AL, E Suarez, K Mehltreter, I Myers-Smith, S Vanderplank, H Slinn, Y Vargas-Rodriguez,
S Haeussler, S David, J Muñoz, R Almazán-Nuñez, D Loughnan, J Benning, D Moeller, J Brodie, H
Thomas, P Morales. 2019.
Science Advances. 5 eaau4403 (more info on project, data, media coverage)
Hargreaves, AL, KE Samis, CG Eckert. 2014.
American Naturalist. 183: 175-183. (blog summary, DOI, Appendices, data)

Species at Risk
Mapping nationally and globally at-risk species to identify hotspots for (and gaps in) conservation
Hardouin, ME, AL Hargreaves. 2023.
Proceedings of the Royal Society B. 290 (1995), 20222307.
High conservation priority of range-edge plant populations not matched by habitat protection or research effort. Caissy, P, S Klemet-N’Guessan, R Jackewi, CG Eckert, AL Hargreaves. 2020.
Biological Conservation. 249: 108732 (data & Rcode)
Spatial distribution and conservation hotspots of mammals in Canada
Cameron, V, AL Hargreaves. 2020.
FACETS. 5: 692-703
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Dispersal
Adaptation to fragmentation: evolutionary dynamics driven by human influences
Cheptou, PO, AL Hargreaves, D Bonte, H Jacquemyn. 2017. Philosophical Transactions of the Royal Society B. 372 (1712), 20160037.
Local maladaptation reduces expansion load during range expansion.
Gilbert K, N Sharp, A Angert*, G Conte*, J Draghi*, F Guillaume*, AL Hargreaves*, R Matthey-Doret*, M Whitlock. 2017. *order alphabetical
American Naturalist. 189: 368-380. (DOI, summary)
Hargreaves, AL, SF Bailey, RA Laird. 2015.
Journal of Evolutionary Biology. 28: 1489-1501. (Appendix, data & R code)
Hargreaves, AL & CG Eckert. 2014.
Functional Ecology. 28: 5-21. (summary)


Pollination & Pollen theft
Narrow entrance of short-tubed Aloe flower facilitates pollen transfer on long sunbird bills.
Hargreaves, AL, GT Langston, SD Johnson. 2019.
South African Journal of Botany. 124: 23-28. (abstract)
High-elevation range limit of an annual herb is neither caused nor reinforced by declining pollinator service. Hargreaves, AL, JL Weiner, CG Eckert. 2015.
Journal of Ecology. 103: 572-584. (PDF, data)
Floral traits mediate the vulnerability of aloes to pollen theft and inefficient pollination by bees.
Hargreaves, AL, LD Harder & SD Johnson. 2012.
Annals of Botany. 109: 761-772. (PDF)
Native pollen thieves reduce the reproductive success of a hermaphroditic plant, Aloe maculata.
Hargreaves, AL, LD Harder & SD Johnson. 2010.
Ecology 91:1693-1703. (PDF)
Consumptive emasculation: the ecological and evolutionary consequences of pollen theft.
Hargreaves, AL, LD Harder & SD Johnson. 2009.
Biological Reviews 84: 259-276. (DOI, PDF)
Aloe inconspicua: the first record of an exclusively insect-pollinated aloe.
Hargreaves, AL, LD Harder & SD Johnson. 2008.
South African Journal of Botany 74: 606-612. (PDF)
Empty flowers as a pollination-enhancement strategy.
Bailey, SF, AL Hargreaves, SD Hechtenthal, RA Laird, TM Latty, TG Reid, AC Teucher & JR Tindall. 2007.
Evolutionary Ecology Research 9: 1245-1262. (PDF)
Dark, bitter-tasting nectar functions as a filter of flower visitors in a bird-pollinated plant.
Johnson, SD, AL Hargreaves & M Brown. 2006.
Ecology 87: 2709-2716. (PDF)
Do floral syndromes predict specialization in plant pollination systems? An experimental test in an "ornithophilous" African Protea.
Hargreaves, AL, SD Johnson & E Nol. 2004.
Oecologia 140: 295-301. (PDF)


Side projects & Miscellaneous
D MacNearney, E Davis, J Rausch, E Nol, G Gilchrist, 15 authors including AL Hargreaves. 2025.
Arctic Science 11, 1-18
An arctic breeding songbird overheats during intense activity even at low air temperatures
O'Connor, RS, OP Love, L Regimbald, A Gerson, K Elliott, AL Hargreaves, F Vézina. 2024.
Scientific Reports. doi.org/10.1038/s41598-024-65208-9
O’Connor, R, A Le Pogam, K Young, O Love, C Cox, G Roy, F Robitaille, K Elliott, AL Hargreaves, E Choy, HG Gilchrist, D Berteaux, A Tam, F Vézina. 2022.
Proceedings of the Royal Society B. Accepted
Limited heat tolerance in a cold-adapted seabird: implications of a warming Arctic
Choy, ES, RS O'Connor, HG Gilchrist, AL Hargreaves, OP Love, F Vézina, KH Elliot. 2021.
Journal of Experimental Biology. 224: jeb242168
O'Connor, Le Pogam, K Young, F Robitaille, E Choy, O Love, K Elliott, A Hargreaves, D Berteaux, A Tam, F Vézina. 2021.
Ecology and Evolution. 11: 1609-1619.
Clustered vs. catastrophic global vertebrate declines
Leung B, AL Hargreaves, D Greenberg, B McGill, M Dornelas, R Freeman. 2020.
Nature. 588: 267-271 (PDF)
Thermal tolerance patterns across latitude & elevation.
Sunday, J, J Bennett, P Calosi, S Clusella-Trullas, S Gravel, AL Hargreaves, F Leiva, et al. 2019.
Philosophical Transactions of the Royal Soc. B. 374.
Concentrations of 17 elements, including mercury, in the tissues, food & abiotic environment of Arctic shorebirds
Hargreaves, AL, DP Whiteside & G Gilchrist. 2011.
Science of the Total Environment. 409: 3757-3770. (DOI, PDF, Appendices)
Concentrations of 17 elements, including mercury, and their relationship to fitness measures in arctic shorebirds and their eggs
Hargreaves, AL, DP Whiteside & G Gilchrist. 2010.
Science of the Total Environment 408: 3153-3161. (DOI, PDF, Appendix)